1. Bousseau S, Brillard JP, Marquant-Le Guienne B, et al. Comparison of bacteriological qualities of various egg yolk sources and the in vitro and in vivo fertilizing potential of bovine semen frozen in egg yolk or lecithin-based diluents. Theriogenology 1998; 50(5): 699-706.
2. Tosic J, Walton A. Effect of egg-yolk and its constituents on the respiration and fertilizing capacity of spermatozoa. J Agric Sci 1947; 37(1): 69-76.
3. Anand M, Yadav S, Vaswani S, et al. Low-density lipoprotein (LDL) as a cryoprotectant in semen extender: A new approach. Asian J Anim Sci 2015; 10: 211-219.
4. Yamauchi S, Nakamura S, Lay KM, et al. Characteristics of Okinawan native agu pig spermatozoa after addition of low-density lipoprotein to freezing extender. J Reprod Dev 2009; 55(5): 558-565.
5. Layek SS, Mohanty TK, Kumaresan A, et al. Cryopreservation of bull semen: evolution from egg yolk based to soybean based extenders. Anim Reprod Sci 2016; 172: 1-9.
6. Lane M, Thérien I, Moreau R, et al. Heparin and high-density lipoprotein mediate bovine sperm capacitation by different mechanisms. Biol Reprod 1999; 60(1): 169-175.
7. Bustani GS, Baiee FH. Semen extenders: an evaluative overview of preservative mechanisms of semen and semen extenders. Vet World 2021; 14(5): 1220-1233.
8. Miguel-Jimenez S, del Alamo MM, Álvarez-Rodríguez M, et al. In vitro assessment of egg yolk-, soya bean lecithin- and liposome-based extenders for cryopreservation of dairy bull semen. Anim Reprod Sci 2020; 215: 106315. doi: 10.1016/j.anireprosci. 2020.106315.
9. Liang J, Larbi A, Lv C, et al. Fertility results after exocervical insemination using goat semen cryopreserved with extenders based on egg yolk, skim milk, or soybean lecithin. Reprod Domest Anim 2023; 58(3): 431-442.
10. Dziekońska A, Neuman NM, Burdal KK, et al. The effect of different extenders on the quality characteristics of European red deer epididymal sperm stored at 5 °C. Animals (Basel) 2022; 12(19): 2669. doi: 10.3390/ ani12192669.
11. Divar MR, Mogheiseh A, Mohammadi F, et al. Effects of extender filtration and egg yolk concentration on canine semen cryopreservation. Reprod Domest Anim 2023; 58(2): 272-287.
12. Hussain Shah SA, Hassan Andrabi SM, Ahmed H, et al. Chicken egg yolk plasma in tris-citric acid extender improves the quality and fertility of cryopreserved water buffalo (Bubalus bubalis) spermatozoa. Theriogenology 2017; 89: 32-40.
13. Moussa M, Martinet V, Trimeche A, et al. Low density lipoproteins extracted from hen egg yolk by an easy method: cryoprotective effect on frozen-thawed bull semen. Theriogenology 2002; 57(6): 1695-1706.
14. Ashrafi I, Daghigh Kia H, Parrish J. Egg yolk saline soluble fraction was as efficient as ammonium sulfate insoluble yolk fraction in cryopreservation of bull semen compared to whole egg yolk. Rev Med Vet 2019; 170(4-6): 104-109.
15. Brito MF, Neves BP, de Oliveira Andrade G, et al. Low density lipoproteins at 2% concentration can replace whole egg yolk in TES-Tris-milk extender for freezing buffalo sperm cells. Acta Sci Vet 2018; 46(1): 6. doi: 10.22456/1679-9216.81815.
16. Gil J, Fierro S, Bentancur O, et al. Chilled storage of ram semen improves with the addition of egg yolk and glycerol to milk‐based extenders. Reprod Domest Anim 2011; 46(3): 503-507.
17. Bahmani S, Eslami M, Farrokhi-Ardabili F, et al. Evaluation of chicken egg yolk Plasma and low-density lipoprotein alone or enriched with ewe or cow skim milk in Tris-citric acid-based diluent for cryostorage of ram semen. Biopreserv Biobank 2023; 21(4): 346-354.
18. Campbell RC, Dott HM, Glover TD. Nigrosin eosin as a stain for differentiating live and dead spermatozoa. J Agric Sci 1956; 48(1): 1-8.
19. Buckett WM, Luckas MJ, Aird IA, et al. The hypo-osmotic swelling test in recurrent miscarriage. Fertil Steril 1997; 68(3): 506-509.
20. Divar MR, Azari M, Mogheiseh A, et al. Supplementation of melatonin to cooling and freezing extenders improves canine spermatozoa quality measures. BMC Vet Res 2022; 18(1): 86. doi: 10.1186/s12917-022-03186-8.
21. Fernández JL, Muriel L, Rivero MT, et al. The sperm chromatin dispersion test: a simple method for the determination of sperm DNA fragmentation. J Androl 2003; 24(1): 59-66.
22. Gharibi S, Niasari-Naslaji A, Poursasan N, et al. Replacement of salamon with shotor diluent and egg yolk with low-density lipoprotein for chilled storage of ram semen. Iran J Vet Res 2014; 15(3): 279-284.
23. Dalal J, Chandolia RK, Pawaria S, et al. Low‐density lipoproteins protect sperm during cryopreservation in buffalo: unraveling mechanism of action. Mol Reprod Dev 2020; 87(12): 1231-1244.
24. Hu JH, Jiang ZL, Lv RK, et al. The advantages of low-density lipoproteins in the cryopreservation of bull semen. Cryobiology 2011; 62(1): 83-87.
25. Dong QX, Rodenburg SE, Hill D, et al. The role of low-density lipoprotein (LDL) and high-density lipoprotein (HDL) in comparison with whole egg yolk for sperm cryopreservation in rhesus monkeys. Asian J Androl 2011; 13(3): 459-464.
26. Wang P, Wang YF, Wang CW, et al. Effects of low- density lipoproteins extracted from different avian yolks on boar spermatozoa quality following freezing-thawing. Zygote 2014; 22(2): 175-181.
27. Bergeron A, Manjunath P. New insights towards understanding the mechanisms of sperm protection by egg yolk and milk. Mol Reprod Dev 2006; 73(10):1338-1344.
28. Caballero I, Parrilla I, Almiñana C, et al. Seminal plasma proteins as modulators of sperm function and their application in sperm biotechnologies. Reprod Domest Anim 2012; 47(Suppl 3): 12-21.
29. Saleh DM, Sumaryadi MY, Nugroho AP, et al. Effect of the addition of egg yolk to skim milk extender and storage time on the motility and fertility of kampung rooster spermatozoa. In proceedings: International conference on improving tropical animal production for food security. Kendari, Indonesia 2022; 101-104.
30. Leboeuf B, Guillouet P, Batellier F, et al. Effect of native phosphocaseinate on the in vitro preservation of fresh semen. Theriogenology 2003; 60(5): 867-877.
31. Šimoník O, Stádník L, Rajmon R, et al. Influence of LDL addition on cryoprotective properties of bovine semen extenders. In proceedings: IV international symposium Agrosym, Jahorina, Bosnia and Herzegovina 2013; 3-6.
32. Yassen AM, Foote RH. Freezability of bull semen in centrifuged or dialyzed and reconstituted buffered egg yolk extenders. J Dairy Sci 1967; 50(9): 1481-1485.
33. García W, Tabarez A, Palomo MJ. Effect of the type of egg yolk, removal of seminal plasma, and donor age on ram sperm cryopreservation. Anim Reprod 2017; 14(4): 1124-1132.
34. Garcia MA, Graham EF. Effects of low-molecular-weight fractions (LMWF) from milk, egg yolk, and seminal plasma on freezability of bovine spermatozoa. Cryobiology 1987; 24(5): 429-436.
35. Fernández-Santos MR, Esteso MC, Montoro V, et al. Cryopreservation of Iberian red deer (Cervus elaphus hispanicus) epididymal spermatozoa: effects of egg yolk, glycerol and cooling rate. Theriogenology 2006; 66(8): 1931-1942.
36. Del Valle I, Souter A, Maxwell WM, et al. Function of ram spermatozoa frozen in diluents supplemented with casein and vegetable oils. Anim Reprod Sci 2013; 138(3-4): 213-219.
37. Susilowati S, Sardjito T, Mustofa I, et al. Effect of green tea extract in extender of Simmental bull semen on pregnancy rate of recipients. Anim Biosci 2020; 34(2): 198-204.
38. Raheja N, Choudhary S, Grewal S, et al. A review on semen extenders and additives used in cattle and buffalo bull semen preservation. J Entomol Zool Stud 2018; 6(3): 239-245.
39. Ari UÇ, Kulaksiz R, Öztürkler Y. Freezability of Tushin ram semen extended with goat or cow milk based extenders. Reprod Domest Anim 2011; 46(6): 975-979.
40. Bustamante Filho IC, Pederzolli CD, Sgaravatti AM., et al. Skim milk-egg yolk-based semen extender compensates for non-enzymatic antioxidant activity loss during equine semen cryopreservation. Anim Reprod 2018; 6(2): 392-399.
