1. Sutili FJ, Gatlin III DM, Heinzmann BM, et al. Plant essential oils as fish diet additives: benefits on fish health and stability in feed. Rev Aquac 2017; 10(3): 716-726.
2. Zeng Z, Zhang S, Wang H, et al. Essential oil and aromatic plants as feed additives in non-ruminant nutrition: a review. J Anim Sci Biotechnol 2015; 6(1): 7. doi: 10.1186/s40104-015-0004-5.
3. Carson CF, Hammer KA. Chemistry and bioactivity of essential oils. In: Thormar H. (Ed). Lipids and essential oils as antimicrobial agents. Oxford, UK: John Wiley & Sons 2011; 203-238.
4. Góra J, Lis A, Kula J, et al. Chemical composition variability of essential oils in the ontogenesis of some plants. Flavour Fragr J 2002; 17(6): 445-451.
5. Sticher O. Essential oils and herbal substances containing essential oil [German]. In: Sticher O, Heilmann J, Zúndorf I, et al, (Eds). Sticker Pharma-cognosie Phytopharmazie. 10th ed. Stuttgart, Germany: Wissenschaftliche Verlagsgesellschaft 2015; 663-740.
6. Swamy MK, Akhtar MS, Sinniah UR. Antimicrobial properties of plant essential oils against human pathogens and their mode of action: an updated review. Evid Based Complement Alternat Med 2016; 2016: 3012462. doi: 10.1155/2016/3012462.
7. Souza CF, Baldissera MD, Baldisserotto B, et al. Essential oils as stress-reducing agents for fish aquaculture: a review. Front Physiol 2019; 10: 785. doi: 10.3389/fphys.2019.00785.
8. Souza CF, Baldissera MD, Salbego J, et al. Physiological responses of Rhamdia quelen (Siluriformes: Heptapteridae) to anesthesia with essential oils from two different chemotypes of Lippia alba. Neotrop Ichthyol 2017; 15(1): e160083. doi: 10.1590/1982-0224-20160083.
9. Baldissera MD, Souza CF, Júnior GB, et al. Melaleuca alternifolia essential oil enhances the non-specific immune system and prevents oxidative damage in Rhamdia quelen experimentally infected by Aeromonas hydrophila: effects on cholinergic and purinergic systems in liver tissue. Fish Shellfish Immunol 2017; 61: 1-8.
10. de Freitas Souza C, Baldissera MD, Bianchini AE, et al. Citral and linalool chemotypes of Lippia alba essential oil as anesthetics for fish: a detailed physiological analysis of side effects during anesthetic recovery in silver catfish (Rhamdia quelen). Fish Physiol Biochem 2018; 44(1): 21-34.
11. Aydın B, Barbas LAL. Sedative and anesthetic properties of essential oils and their active compounds in fish: a review. Aquaculture 2020; 520: 734999. doi: 10.1016/j.aquaculture.2020.734999.
12. Tarkhani R, Imani A, Jamali H, et al. Anaesthetic efficacy of eugenol on various size classes of angelfish (Pterophyllum scalare Schultze, 1823). Aquac Res 2017; 48(10): 5263-5270.
13. Tarkhani R, Imani A, Jamali H, et al. Anaesthetic efficacy of eugenol on Flowerhorn (Amphilophus labiatus × Amphilophus trimaculatus). Aquac Res 2017; 48(6): 3207-3215.
14. Rombout JH, Abelli L, Picchietti S, et al. Teleost intestinal immunology. Fish Shellfish Immunol 2011; 31(5): 616-626.
15. Ley RE, Lozupone CA, Hamady M, et al. Worlds within worlds: evolution of the vertebrate gut microbiota. Nat Rev Microbiol 2008; 6(10): 776-788.
16. Laparra JM, Sanz Y. Interactions of gut microbiota with functional food components and nutraceuticals. Pharmacol Res 2010; 61(3): 219-225.
17. Nazzaro F, Fratianni F, De Martino L, et al. Effect of essential oils on pathogenic bacteria. Pharmaceuticals (Basel) 2013; 6(12): 1451-1474.
18. Plésiat P, Nikaido H. Outer membranes of Gram‐negative bacteria are permeable to steroid probes. Mol Microbiol 1992; 6(10): 1323-1333.
19. Prasch S, Bucar F. Plant derived inhibitors of bacterial efflux pumps: an update. Phytochem Rev 2015; 14: 961-974.
20. Khalafalaa M, Shehab SM, Aboraya MH, et al. Herbal essential oils improve growth, antioxidant response, and gene expression in Nile Tilapia fingerlings. Front Vet Sci 2025; 12: 1620632. doi: 10.3389/fvets. 2025.1620632.
21. Khorshidi Z, Sarvi Moghanlou K, Imani A, et al. The interactive effect of dietary curcumin and silver nano-particles on gut microbiota of common carp (Cyprinus carpio). Iran J Sci Technol Trans Sci 2016; 42(2): 379-387.
22. Ran C, Hu J, Liu W, et al. Thymol and carvacrol affect hybrid tilapia through the combination of direct stimulation and an intestinal microbiota-mediated effect: insights from a germ-free zebrafish model. J Nutr 2016; 146(5): 1132-1140.
23. Thapa D, Louis P, Losa R, et al. Essential oils have different effects on human pathogenic and commensal bacteria in mixed faecal fermentations compared with pure cultures. Microbiology (Reading) 2015; 161(Pt 2): 441-449.
24. Zhang Y, Wang QC, Yu H, et al. Evaluation of alginate-whey protein microcapsules for intestinal delivery of lipophilic compounds in pigs. J Sci Food Agric 2016; 96(8): 2674-2681.
25. Mohamed GA, Amhamed ID, Almabrok AA, et al. Effectof celery (Apium graveolens) extract on the growth, haematology, immune response and digestive enzyme activity of common carp (Cyprinus carpio). Mar Sci Technol Bull 2018; 7(2): 51-59.
26. Ghafarifarsani H, Kachuei R, Imani A. Dietary supplementation of garden thyme essential oil ameliorated the deteriorative effects of aflatoxin B1 on growth performance and intestinal inflammatory status of rainbow trout (Oncorhynchus mykiss). Aquaculture 2021; 531: 735928. doi: 10.1016/ j.aquaculture.2020.735928.
27. Tafi AA, Meshkini S, Tukmechi A, et al. Therapeutic and histopathological effect of Aloe vera and Salvia officinalis hydroethanolic extracts against Streptococcus iniae in rainbow trout. Arch Razi Inst 2020; 75(2): 257-287.
28. Imani A, Salimi Bani M, Noori F, et al. The effect of bentonite and yeast cell wall along with cinnamon oil on aflatoxicosis in rainbow trout (Oncorhynchus mykiss): digestive enzymes, growth indices, nutritional performance and proximate body composition. Aquaculture 2017; 476: 160-167.
29. Khani S, Sarvi Moghanlou K, Imani A, et al. The protective effect of dietary cinnamon essential oil (Cinnamomum verum) supplementation in reducing the toxicity of aflatoxin B1 to rainbow trout (Oncorhynchus mykiss) fingerlings. J Fish 2017; 69(4): 481-495.
30. Salem MOA. Effects of plant essential oil used as feed supplements in aquaculture on immune response, and antioxidant status of finfish: a review. JMSET 2023; 9(1): E12-E40.
31. Tort L. Stress and immune modulation in fish. Dev Comp Immunol 2011; 35(12): 1366-1375.
32. Biller JD, Takahashi LS. Oxidative stress and fish immune system: phagocytosis and leukocyte respiratory burst activity. An Acad Bras Cienc 2018; 90(4): 3403-3414.
33. Birnie-Gauvin K, Costantini D, Cooke SJ, et al. A comparative and evolutionary approach to oxidative stress in fish: a review. Fish Fish 2017; 18(5): 928-942.
34. Dawood MA, El Basuini MF, Yilmaz S, et al. Exploring the roles of dietary herbal essential oils in aquaculture: a review. Animals 2022; 12(7): 823. doi: 10.3390/ ani12070823.
35. Zhang R, Wang XW, Liu LL, et al. Dietary oregano essential oil improved the immune response, activity of digestive enzymes, and intestinal microbiota of the koi carp, Cyprinus carpio. Aquaculture 2020; 518: 734781. doi: 10.1016/j.aquaculture.2019. 734781.
36. El-Hawarry WN, Mohamed RA, Ibrahim SA. Collaborating effects of rearing density and oregano oil supplementation on growth, behavioral and stress response of Nile tilapia (Oreochromis niloticus). Egypt J
Aquat Res 2018; 44(2): 173-178.
37. Giannenas I, Triantafillou El, Stavrakakis S, et al. Assessment of dietary supplementation with carvacrol or thymol containing feed additives on performance, intestinal microbiota and antioxidant status of rainbow trout (Oncorhynchus mykiss). Aquaculture 2012; 350-353: 26-32.
38. Peterson BC, Bosworth BG, Li MH, et al. Assessment of a phytogenic feed additive (Digestarom PEP MGE) on growth performance, processing yield, fillet composition, and survival of channel catfish. J World Aquac Soc 2014; 45(2): 206-212.
39. Zeppenfeld CC, Saccol EMH, Pês TS, et al. Aloysia triphylla essential oil as food additive for Rhamdia quelen - Stress and antioxidant parameters. Aquac Nutr 2017; 23(6): 1362-1367.
40. Hsieh TJ, Wang JC, Hu CY, et al. Effects of rutin from Toona sinensis on the immune and physiological responses of white shrimp (Litopenaeus vannamei) under Vibrio alginolyticus challenge. Fish Shellfish Immunol 2008; 25(5): 581-588.
41. Aydın B, Akhan S, Gümüş E, et al. Anesthetic efficacy of clove oil and 2- phenoxyethanol on doctor fish, Garra rufa (Heckel, 1843). Bol Inst Pesca 2019; 45(4): e506. doi: 10.20950/1678-2305.2019. 45.4.506.
42. Metin S, Didinen BI, Kubilay A, et al. Determination of anesthetic effects of some medicinal plants on rainbow trout (Oncorhynchus mykiss Walbaum, 1792). LimnoFish 2015; 1(1): 37-42.
43. Saccol EMH, Uczay J, Pês TS, et al. Addition of Lippia alba (Mill) N. E. Brown essential oil to the diet of the silver catfish: an analysis of growth, metabolic and blood parameters and the antioxidant response. Aquaculture 2013; 416-417: 244-254.
44. Barbas LAL, Hamoy M, de Mello VJ, et al. Essential oil of citronella modulates electrophysiological responses in tambaqui Colossoma macropomum: a new anaesthetic for use in fish. Aquaculture 2017; 479: 60-68.
45. Baldissera MD, Souza CF, Baldisserotto B. Melaleuca alternifolia essential oil prevents bioenergetics dysfunction in spleen of silver catfish naturally infected with Ichthyophthirius multifiliis. Microb Pathog 2018; 123: 47-51.
46. Tahri Mirghaed A, Ghelichpour M, Hoseini SM. Myrcene and linalool as new anesthetic and sedative agents in common carp, Cyprinus carpio - comparison with eugenol. Aquaculture 2016; 464: 165-170.
47. Yousefi M, Hoseinifar SH, Ghelichpour M, et al. Anesthetic efficacy and biochemical effects of citronellal and linalool in common carp (Cyprinus carpio Linnaeus, 1758) juveniles. Aquaculture 2018; 493: 107-112.
48. Mazandarani M, Hoseini SM. Menthol and 1, 8‐cineole as new anaesthetics in common carp, Cyprinus carpio (Linnaeus, 1758). Aquac Res 2016; 48(6): 3041-3051.
49. Baldissera MD, Souza CF, Zeppenfeld CC, et al. Dietary supplementation with nerolidol nanospheres improves growth, antioxidant status and fillet fatty acid profiles in Nile tilapia: benefits of nanotechnology for fish health and meat quality. Aquaculture 2020; 516: 734635. doi: 10.1016/ j.aquaculture.2019.734635.
50. Yostawonkul J, Kitiyodom S, Kaewmalun S, et al. Bifunctional clove oil nanoparticles for anesthesia and anti-bacterial activity in Nile tilapia (Oreochromis niloticus). Aquaculture 2019; 503: 589-595.
51. Kheawfu K, Pikulkaew S, Chaisri W, et al. Nanoemulsion: a suitable nanodelivery system of clove oil for anesthetizing Nile tilapia. Drug Discov Ther 2017; 11(4): 181-185.
52. Kheawfu K, Pikulkaew S, Rades T, et al. Development and characterization of clove oil nanoemulsions and self-microemulsifying drug delivery systems. J Drug Deliv Sci Technol 2018; 46: 330-338.
53. Pierozan MB, Oliveira Filho JG, Cappato LP, et al. Essential oils against spoilage in fish and seafood: impact on product quality and future challenges. Foods 2024; 13(23): 3903. doi: 10.3390/foods13233903.
54. Farhadi N, Meshkini S, Mehdizadeh T. Effect of chitosan edible coating containing Froriepia subpinnata extract on the shelf life of Nile Tilapia (Oreochromis niloticus) fillet at refrigerated temperature. JRIFST 2022; 11(1): 95-108.
55. Saeid Asr E, Naghibi S, Mojaddar Langroodi A, et al. Impact of carboxymethyl cellulose coating incorporated with rosemary essential oil and sodium acetate on the quality and shelf life of rainbow trout fillet. J Aquat Food Prod Technol 2021; 30(1): 16-30.
56. Kolygas MN, Bitchava K, Nathanailides C, et al. Phytochemicals: essential oils and other extracts for disease prevention and growth enhancement in aquaculture: challenges and opportunities. Animals (Basel) 2025; 15(18): 2653. doi: 10.3390/ ani15182653.
57. Soengas JL, Cerdá-Reverter JM, Delgado MJ. Central regulation of food intake in fish: an evolutionary perspective. J Mol Endocrinol 2018; 60(4): R171-R199.
58. Benomari FZ, Sarazin M, Chaib D, et al. Chemical variability and chemotype concept of essential oils from Algerian wild plants. Molecules 2023; 28(11): 4439. doi: 10.3390/molecules28114439.
59. Hassan HU, Ali A, Al Sulivany BSA, et al. Investigation of the effects of phytogenic dietary additives on growth performance, nutrient utilization, economic efficiency and health of Pangasius hypophthalmus: implications for sustainable aquaculture development. Sci Rep 2025; 15(1): 22661. doi: 10.1038/s41598-025-03658-5.